Sites, Jack

Biology

Email: 3MdfnbVlwhvCe|x1hgx  (Email Form)

Website: http://siteslab.byu.edu/

4029  LSB
Provo, UT 84602
(801) 422-2279

Professor

 Teaching Interests
I have taught the following classes/seminars at BYU:
Biol. 100 (Life Sciences for Non-majors)
Biol. 220A (Animal Diversity [vertebrate section only])
Biol. 220 (Biodiversity)
Zool. 363 (Comparative Vertebrate Anatomy)
Zoo?Bioll. 445 (Herpetology)
Zool/Biol. 470 (Conservation Biology)
Zool/Biol. 475 (Evolutionary Biology)
Zool. 549R (Advanced Topics in Evolutionary Genetics)
Zool. 559R (seminars in Conservation Biology, Molecular Evolution, Speciation;
494R field courses: Biodiversity of Appalachia [1999], Tropical Biology-Amazonia [2000]; Tropical Biology - French Guiana [2006]
Zool. 604 (Phylogenetic Systematics)
Zool. 605 (Molecular Methods in Systematics & Population Biology)
 Research Interests
Herpetology, Speciation & Species Delimitation, Hybrid Zone Dynamics, Origins of Parthenogenesis, Phylogeography, Systematic Biology, Conservation & Population Genetics, Biodiversity, Public Outreach
 Memberships
  • National Center for Science Education, 2002-Present
  • Chelonian Research Foundation, 1996-Present
  • American Association for the Advancement of Science, 1990-Present
  • Bean Museum, 1982-Present
  • American Society of Ichthyologists and Herpetologists, 1975-Present
  • Society for the Study of Amphibians and Reptiles, 1975-Present
  • Society of Conservation Biology, 1980-2009
 Honors and Awards
  • Society of Systematic Biologists (SSB) : President Elect
  • College of Life Sciences : Thomas L. Martin Professorship
  • Brigham Young University : Jessie Knight University Professorship
  • NSF, MEG, National Geographic : see research profile
 Courses Taught

Summer 2016
  • BIO 799R: Dissertation Section 002
Spring 2016
  • BIO 799R: Dissertation Section 002
Winter 2016
  • BIO 445: Herpetology Section 001
  • BIO 494R: Mentored Research Section 021
  • BIO 559R: Adv Topics Ecology & Evolution Section 002
  • BIO 799R: Dissertation Section 016
Fall 2015
  • BIO 220: Biological Diversity: Animals Section 001
  • BIO 450: Conservation Biology Section 001
  • BIO 494R: Mentored Research Section 020
  • BIO 559R: Adv Topics Ecology & Evolution Section 002
  • BIO 799R: Dissertation Section 002

Selected Publications

Journal Articles

Breitman MF, Bonino M, Sites, Jr. JW, Avila LJ, Morando M. 2015. Morphological variation, niche divergence, and phylogeography of lizards of the Liolaemus lineomaculatus section (Liolaemini) from Southern Patagonia. Herpetological Monographs. 29:65-88.

Morando M, Olave M, Avila LJ, Baker E, Sites, Jr. JW. 2015. Molecular phylogeny of the lizard clade Leiosaurae endemic to southern South America. Herpetologica. 71:322-331.

Aguilar C, Stark MR, Arroyo J, Standing MD, Rios S, Washburn T, Sites, Jr. JW. 2015. Placental morphology in two sympatric Andean lizards of the genus Liolaemus (Reptilia: Liolaemidae). Journal of Morphology. 276::1205-1217.

Breitman MF, Ramiro JN, Avila LJ, Sites, Jr. JW, Morando M. 2015. Phylogeography and morphological variation of the northernmost distributed species of the Liolaemus lineomaculatus section (Liolaemini) from Patagonia. Amphibia-Reptilia. 36:373-387.

Grismer LL, Wood, Jr. PL, Lee CH, Quah ES, Anuar S, Ngadi E, Sites, Jr. JW. 2015. An integrative taxonomic review of the agamid genus Bronchocela (Kulh, 1820) from Peninsular Malaysia with descriptions of new montane and insular endemics. Zootaxa. 3848:1-23.

Olave M, Avila LJ, Sites, Jr. JW, Morando M. 2015. Model-based approach to test hard polytomies in the Eulaemus clade of the most diverse South American lizard genus Liolaemus (Liolaemini, Squamata). Zoological Journal of the Linnaean Society. 174:169-184.

Avila LJ, Medina C, Perez C, Sites, Jr. JW, Morando M. 2015. Molecular phylogenetic relationships of the lizard clade Liolaemus elongatus (Iguania: Liolaemini) with a description of a new species from an isolated volcanic peak in northern Patagonia. Zootaxa. 3947:67-84.

Medina C, Avila LJ, Sites, Jr. JW, Morando M. 2015. Molecular phylogeny of the Liolaemus kriegi complex (Iguania, Liolaemini). Herpetologica. 71:143-151.

Reeder TW, Townsend T, Mulcahy D, Noonan B, Wood P, Sites, Jr. JW, Wiens J. 2015. Integrated analyses resolve conflicts over squamate reptile phylogeny and reveal unexpected placements of fossil taxa. PLoS ONE. 10(3)(e0118199). <website> doi:10.1371/journal.pone

Breitman MF, Minoli I, Avila LJ, Medina CD, Sites, Jr. JW, Morando M. 2014. Lagartijas de la provincia de Santa Cruz, Argentina: distribution geográfica, diversidad genética y estado de conservación. Cuadernos de Herpetologia. 28:83-110.

Recoder RS, Werneck FP, Teixeira, Jr. M, Colli GR, Sites, Jr. JW, Rodrigues MT. 2014. Geographic variation and systematic review of the lizard genus Vanzosaura (Squamata, Gymnophthalmidae), with the description of a new species. Zoological Journal of the Linnean Society. 171:206-225.

Olave M, Avila LJ, Sites, Jr. JW, Morando M. 2014. Multilocus phylogeny of the widely distributed South American lizard clade Eulaemus (Liolaemini, Liolaemus). Zoological Scripta. 43:323-337.

Medina CD, Avila LJ, Sites, Jr. JW, Morando M. 2014. Multilocus phylogeography of the Patagonian lizard complex Liolaemus kriegi (Iguania: Liolaemini). Biological Journal of the Linnean Society. 113:256-269.

Breitman F, Avila LJ, Parra M, Sites JW, Morando M. 2012. How lizards survived blizzards: phylogeograhy of the Liolaemus lineomaculatus group (Liolaemidae) reveals multiple breaks and refugia in southern Patagonia, and their concordance with other co-distributed taxa. Molecular Ecology. 21:6068-6085.

Hamilton BT, Hart S, Sites JW. 2012. Feeding ecology of the milksnake (Lampropeltis triangulum, Colubridae) in the western United States. Journal of Herpetology. 46:515-522.

Vera-Escalona I, D'Elia G, Gouin N, Fontanella FM, Munoz-Mendoza C, Sites JW, Victoriano PF. 2012. Lizards on ice; evidence for multiple refugia in Liolaemus pictus (Squamata: Liolaemidae) during the Last Glacial Maximum in the Southern Andean Beech Forests. PLoS ONE. 7(11): e48358. doi:10.1371/journal.pone.0048358.

Mulcahy DG, Noonan BP, Moss T, Townsend TM, Reeder TW, Sites JW, Wiens JJ. 2012. Comparing the performance of dating methods using phylogenomic data in squamate reptiles. Molecular Phylogenetics & Evolution. 65:974-994.

Werneck FP, Gamble T, Colli GR, Rodrigues MT, Sites JW. 2012. Deep diversification and long-term persistence in the South American ‘dry diagonal’: integrating continent-wide phylogeography and distribution modeling of geckos. Evolution. 66:3014-3034.

Werneck FP, Nogueira C, Colli GR, Sites JW, Costa GC. 2012. Climatic stability in the Brazilian Cerrado: implications for biogeographical connections of the South American savannas, species richness, and conservation in a biodiversity hotspot. Journal of Biogeography. 39:1695-1706.

Avile-Pires TC, Mulcahy DG, Werneck FP, Sites JW. 2012. Phylogeography of Kentropyx calcarata and Gonatodes humeralis (Reptilia: Squamata) in eastern Amazonia, Brazil. Herpetologica. 68:272-287.

Wiens JJ, Hutter CR, Mulcahy DG, Noonan BP, Townsend TM, Sites JW, Reeder TW. 2012. Resolving the phylogeny of lizards and snakes with extensive sampling of genes and species. Biology Letters. doi:10.1098/rsbl.2012.0703.

Camargo A, Morando M, Avila LJ, Sites JW. 2012. Species delimitation with ABC and other coalescent-based methods in lizards of the Liolaemus darwinii complex (Squamata: Liolaemidae). Evolution. 66:2834-2849.

Avila LJ, Perez CH, Medina CD, Sites JW, Morando M. 2012. A new species lizard of the Liolaemus elongatus clade (Squamata: Iguania: Liolaemini) from Curi Leuvu River Valley, northern Patagonia, Neuquén, Argentina. Zootaxa. 3325:37-52.

Camargo A, Avila LJ, Morando M, Sites JW. 2012. Accuracy and precision in species tree estimation: an empirical evaluation of performance in lizards of the Liolaemus darwinii group (Squamata: Liolaemidae) under varying sub-sampling designs. Systematic Biology. 61:272-288.

Sites JW. 2012. Wilmer Webster Tanner: 17 December 1909 – 28 October 2011. Herpetological Review. 43:12-15.

Fontanella FM, Feltrin N, Avila LJ, Sites JW, Morando M. 2012. Early stages of divergence: phylogeography, climate modeling, and niche differentiation in the South American lizard Liolaemus petrophilus (Squamata: Liolaemidae). Ecology and Evolution. 2:792-808.

Hannah N, Brown D, Avila LJ, Sites JW, Morando M, Fontanella FM. 2012. Characterization of 10 polymorphic microsatellite loci in the South American lizard Liolaemus fitsingerii with cross-amplifiction in L. chehuachekenk. Conservation Genetics Resources. 4:105-107.

Mendelson JR, Mulcahy DG, Williams TS, Sites JW. 2011. A phylogeny and evolutionary natural history of Mesoamerican toads (Anura: Bufonidae: Incilius). Zootaxa. 3138:1-34.

Sites JW, Avila LJ. 2010. A new species of the Liolaemus elongatus group (Squamata: Iguania: Liolaemini) from Cordillera del Veinto, northwestern Patagonia, Neuquen, Argentina. Zootaxa. 2667:28-42.

Wiens JJ, Kuczynski CA, Townsend T, Reeder TW, Mulcahy D, Sites JW. 2010. Combining phylogenomics and fossils in higher-level squamate phylogeny: molecular data change the placement of fossil taxa. Systematic Biology. 59(6):674-688. <website>

Camargo A, Sinervo BR, Sites JW. 2010. Lizards as model organisms for linking phylogeographic and speciation studies. Molecular Ecology. 19(16):3250-3270. <website>

Sinclair EA, Pramuk J, Bezy RL, Crandall KA, Sites JW. 2010. DNA evidence for non-hybrid origins of parthenogensis in natural populations of vertebrates. Evolution. 64(5):1346-1357.

Sinervo B, Méndez-de-la-Cruz F, Miles DB, Heulin B, Bastiaans E, Villagrán-Santa Cruz M, Lara-Resendiz R, Martínez-Méndez N, Calderón-Espinosa ML, Meza-Lázaro RN, et al. 2010. Erosion of lizard diversity by climate change and altered thermal niches. Science. 328(5980):894-899. <website>

Avila LJ, Perez C, Morando M, Sites JW. 2010. A new species of Liolaemus (Reptilia: Squamata) from southwestern Rio Negro province, northern Patagonia, Argentina. Zootaxa. 2434:47-59. <website>

Benavides E, Baum R, Snell HM, Snell HL, Sites JW. 2009. Island biogeography of Galapagos lave lizards (Tropiduridae: Microlophus): species diversity and colonization of the archipelago. Evolution. 63:1606-1626.

Sites JW, Marshall JC. 2003. Species delimitation: a Renaissance issue in systematic biology. Trends in Ecology and Evolution. 18::462-470.

Presentations

Camargo A, Morando M, Sites JW, Avila LJ. Evolución morphológico en el grupo Liolaemus darwinii (Squamata: Liolaemidae) sugiere diversificación adaptative asociada a nichos climáticos. II Congreso Uruguayo de Zoología. Montevideo, UR. 2012 .

Camargo A, Werneck F, Morando M, Sites JW, Avila LJ. Expansión de rango, dinámica demográphica y paleo-distribución de Liolaemus darwinii (Squamata: Liolaemidae) durante el Cuaternario del Argentina Central. II Congreso Uruguayo de Zoología. Montevideo, UR. 2012 .

Camargo A, Morando M, Sites JW, Avila LJ. Evolución morphológico en el grupo Liolaemus darwinii (Squamata: Liolaemidae) sugiere diversificación adaptative asociada a nichos climáticos. XIII Congreso Argentina Herpetologia. Mar de La Plata, ARG. 2012 .

Camargo A, Werneck F, Morando M, Sites JW, Avila LJ. Expansión de rango, dinámica demográphica y paleo-distribución de Liolaemus darwinii (Squamata: Liolaemidae) durante el Cuaternario del Argentina Central. XIII Congreso Argentina Herpetologia. Mar de La Plata, ARG. 2012 .

Victoriano P, Vera-Escalona I, Coronado T, Muñoz-Mendoza C, Sites JW. Tendencias geográficas de valores de conservación intraespecíficos en lagartijas del centro-sur de Chile. PIRE-Patagonia meeting: “Conservación en Patagonia: Integrando Ciencia y Gestión”. Bariloche, Argentina. 2012 .

Olave M, Avila L, Sites JW, Morando M. A well-supported species tree of the Eulaemus clade (Squamata: Liolaemus): evidence of an ancient radiation. VII World Congress of Herpetology. Vancouver, BC. 2012 .

Breitman MF, Avila LJ, Parra M, Sites JW, Morando M. How lizards survived blizzards: phylogeography of the Liolaemus lineomaculatus group (Liolaemini) reveals multiple breaks and refugia in southern Patagonia, and their concordance with other co-distributed taxa. VII World Congress of Herpetology. Vancouver, BC. 2012 .

Morando M, Avila LJ, Perez CH, Hawkins M, Sites JW. How many Phymaturus (Squamata, Liolaemini) are there? A molecular assessment of flat lizards on rock outcrops. VII World Congress of Herpetology. Vancouver, B.C., Canada. 2012 .

Werneck FP, Gamble T, Colli G, Rodrigues MT, Sites JW. Lizard diversification in the South American dry diagonal. VII World Congress of Herpetology. Vancouver, BC. 2012 .

Morando M, Avila LJ, Dawson A, Perez C, Sites JW. Molecular phylogeny of Homonota (Phyllodactylidae): yet another cryptically diverse gecko clade. VII World Congress of Herpetology. Vancouver, BC. 2012 .

Morando M, Avila LJ, Baker E, Sites JW. Molecular phylogeny of the endemic South American lizard clade Leiosaurae. VII World Congress of Herpetology. Vancouver, B.C., Canada. 2012 .

Camargo A, Morando M, Avila LJ, Sites JW. Species delimitation with ABC and other coalescent-based methods. VII World Congress of Herpetology. Vancouver, BC. 2012 .

Olave M, Avila LJ, Sites JW, Morando M. Statistical tests for an ancient radiation of the most diverse reptile genus in South America. 1st Joint Congress of Evolutionary Biology. Ottawa, Ontario, Canada. 2012 .

Sites JW. Species Delimitation: A Decade After the Renaissance. invited research seminar. Brasilia, BRAZIL. 2012 .

Sites JW. Species Delimitation: the Other Goal of Systematic Biology.". invited research seminar. Brasilia, BRAZIL. 2012 .

Brown D, Sites JW, Fontanella FM. A multi-locus phylogenetic analysis of the North American xenodontine snakes. National Conference for Undergraduate Research. Ogden, Ut. 2012 .

Baker E, Morando M, Avila LJ, Sites JW, Fontanella FM. Phylogenetic analysis of the Squamate clade Leiosauridae. National Conference for Undergraduate Research. Ogden, Ut. 2012 .

Victoriano P, Gouin N, D'Elia G, Coronado T, Munoz C, Vera I, Sites JW. Comparing phylogeographies of lizards from the Chilean Biodiversity Hotspot: associations with glaciations and landscape structure. Chilean Evolution Society - annual meeting. Concepcion. 2011 .

Munoz C, Gouin N, D'Elia G, Sites JW, Victoriano P. Phylogeographic comparisons of two Liolaemus from south-central Chile suggest discordant micro-evolutionary processes. Chilean Evolution Society - annual meeting. Concepcion. 2011 .

Sites JW. Phylogenetic Insights on Evolutionary Novelties in Lizards and Snakes. EcoLunch. BYU-Provo. 2011 .

Perez CH, Avila LJ, Olave M, Perez DR, Sites JW, Morando M. A new species of Liolaemus (Iguania: Liolaemini) from a volcanic mountain of northern Neuquen Province, Argentina. XII Congreso Argentino de Herpetología. Bariloche, Argentina. 2011 .

Camargo A, Avila LJ, Morando M, Sites JW. Accuracy and precision of species trees: the effects of locus, individual, and base-pair sampling on inference of species trees in the Liolaemus darwinii group (Squamata, Liolaemidae). XII Congreso Argentino de Herpetología. Bariloche, Argentina. 2011 .

Breitman MF, Parra M, Avila LJ, Sites JW, Morando M. Descrubriendo patrones genéticos y morfológicos de la Patagonia Austral: Lagartijas del grupo Lineomaculatus (Squamata, Iguania, Liolaemini, Liolaemus). XII Congreso Argentino de Herpetología. Bariloche, Argentina. 2011 .

Ferraras I, Olave M, Avila LJ, Sites JW, Morando M. Estudio de la variación fenotípica en los complejos de lagartijas boulengeri y rothi (Squamata: Liolaemus). XII Congreso Argentino de Herpetología. Bariloche, Argentina. 2011 .

Olave M, Avila LJ, Sites JW, Morando M. Filogenia multilocus del clado Eulaemus (Squamata: Liolaemus): Evidencias de division de linaje incomplete, hibridizacion y nuevas species candidatas. XII Congreso Argentino de Herpetología. Bariloche, Argentina. 2011 .

Medina CD, Avila LJ, Perez CH, Perez DR, Sites JW, Morando M. New species of lizards of the Liolaemus elongatus species group (Iguania: Liolaemini) from northwestern Patagonia, Neuquen Province, Argentina. XII Congreso Argentino de Herpetología. Bariloche, Argentina. 2011 .

Camargo A, Morando M, Avila LJ, Sites JW. Species delimitation with ABC and other coalescent-based methods of lizards in the Liolaemus darwinii complex (Squamata, Liolaemidae). XII Congreso Argentino de Herpetología. Bariloche, Argentina. 2011 .

Breitman MF, Perez CH, Parra M, Sites JW. Tres nuevas species de la sección Liolaemus lineomaculatus (Squamata: Sauria: Liolaemidae) de Patagonia. XII Congreso Argentino de Herpetología. Bariloche, Argentina. 2011 .

Aguilar C, Sites JW. A provisional assessment of species limits between two Andean populations of Liolaemus from the montanus group in southern Peru (Squamata; Liolaemidae). IX Latin American Congress of Herpetology. Curitiba, Brazil. 2011 .

Avila-Pires LJ, Medina C, Perez C, Perez D, Sites JW, Morando M. New species of lizards previously confused with Liolaemus elongatus Koslowsky 1896 (Reptilia: Squamata: Iguania: Liolaemini) from Neuquen Province, Argentina. IX Latin American Congress of Herpetology X. Curitiba, Brazil. 2011 .

Sites JW. Rock-Paper-Scissors, Viviparity, and Speciation: Links Between Climate Warming and Lizard Extinctions. Invited Seminar Speaker. Department of Biology, Weber State University. 2011 .

Hannah N, Morando M, Avila LJ, Sites JW, Fontanella F. A microsatellite library of the South American genus Liolaemus: provisional tests of their utility for evolutionary studies. College of Life Sciences: Undergraduate Research Symposium. BYU-Provo. 2011 .

Brown D, Sites JW, Fontanella F. Multi-locus phylogenetic analysis of the North American xenodontine snakes. College of Life Sciences: Undergraduate Research Symposium. BYU-Provo. 2011 .

Baker E, Morando M, Avila LJ, Sites JW, Fontanella F. New insights into phylogenetic relationships and species diversity of the South American lizard species complex Liolaemus fitzingerii, based on combined analysis of mitochondrial and nuclear data. College of Life Sciences: Undergraduate Research Symposium. BYU-Provo. 2011 .

Sites JW. Rock-Paper-Scissors, Viviparity, and Speciation: Links Between Climate Warming and Lizard Extinctions. Cross-program Seminar Speaker. Center for the Environment/Discovery Center, Purdue University. 2011 .

Sites JW. Rock-Paper-Scissors, Viviparity, and Speciation: Links Between Climate Warming and Lizard Extinctions. EcoLunch. BYU-Provo. 2011 .

Technical Report

Minoli I, Morando M, Breitman F, Medina C, Sites JW, Avila LJ. 2012. "Lagartijas de la Provincia de Chubut (Argentina): Distribución Geográfica, Diversidad Genética y Comentarios Acerca de su Estado de Conservación. Informe Técnico .".

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