The College of Life Sciences

Sites, Jack

Biology

Email: 7QhjrfZp{lzGi€|5lk|  (Email Form)

Website: http://siteslab.byu.edu/

695  WIDB
Provo, UT 84602
(801) 422-2279

Professor

 Teaching Interests
I have taught the following classes/seminars at BYU:
Biol. 100 (Life Sciences for Non-majors)
Biol. 220A (Animal Diversity [vertebrate section only])
Biol. 220 (Biodiversity)
Zool. 363 (Comparative Vertebrate Anatomy)
Zoo?Bioll. 445 (Herpetology)
Zool/Biol. 470 (Conservation Biology)
Zool/Biol. 475 (Evolutionary Biology)
Zool. 549R (Advanced Topics in Evolutionary Genetics)
Zool. 559R (seminars in Conservation Biology, Molecular Evolution, Speciation;
494R field courses: Biodiversity of Appalachia [1999], Tropical Biology-Amazonia [2000]; Tropical Biology - French Guiana [2006]
Zool. 604 (Phylogenetic Systematics)
Zool. 605 (Molecular Methods in Systematics & Population Biology)
 Research Interests
Herpetology, Speciation & Species Delimitation, Hybrid Zone Dynamics, Origins of Parthenogenesis, Phylogeography, Systematic Biology, Conservation & Population Genetics, Biodiversity, Public Outreach
 Memberships
  • National Center for Science Education, 2002-Present
  • Chelonian Research Foundation, 1996-Present
  • American Association for the Advancement of Science, 1990-Present
  • Bean Museum, 1982-Present
  • American Society of Ichthyologists and Herpetologists, 1975-Present
  • Society for the Study of Amphibians and Reptiles, 1975-Present
  • Society of Conservation Biology, 1980-2009
 Honors and Awards
  • Society of Systematic Biologists (SSB) : President Elect
  • College of Life Sciences : Thomas L. Martin Professorship
  • Brigham Young University : Jessie Knight University Professorship
  • NSF, MEG, National Geographic : see research profile
 Courses Taught

Fall 2013
  • BIO 220A: Biological Diversity: Animals Section 001
  • BIO 450: Conservation Biology Section 001
  • BIO 559R: Adv Topics Ecology & Evolution Section 002
  • BIO 799R: Dissertation Section 002
Summer 2013
  • BIO 799R: Dissertation Section 002
Spring 2013
  • BIO 799R: Dissertation Section 002
Winter 2013
  • BIO 494R: Mentored Research Section 004
  • BIO 559R: Adv Topics Ecology & Evolution Section 006
  • BIO 799R: Dissertation Section 001

Selected Publications
Journal Articles

Aguilar C, Wood, Jr. PL, Cusi JC, Guzman A, Huari F, Lundberg M, Mortensen E, Ramirez C, Robles D, Suarez J, et al. Integrative taxonomy and preliminary assessment of species limits in the Liolaemus walkeri complex (Squamata, Liolaemidae), with descriptions of three new species from Peru. ZooKeys. 2013 Oct;364:47 - 91.

Breitman F, Avila LJ, Parra M, Sites JW, Morando M. How lizards survived blizzards: phylogeograhy of the Liolaemus lineomaculatus group (Liolaemidae) reveals multiple breaks and refugia in southern Patagonia, and their concordance with other co-distributed taxa. Molecular Ecology. 2012 Dec;21:6068-6085.

Hamilton BT, Hart S, Sites JW. Feeding ecology of the milksnake (Lampropeltis triangulum, Colubridae) in the western United States. Journal of Herpetology. 2012 Dec;46:515-522.

Vera-Escalona I, D'Elia G, Gouin N, Fontanella FM, Munoz-Mendoza C, Sites JW, Victoriano PF. Lizards on ice; evidence for multiple refugia in Liolaemus pictus (Squamata: Liolaemidae) during the Last Glacial Maximum in the Southern Andean Beech Forests. PLoS ONE. 2012 Nov:7(11): e48358. doi:10.1371/journal.pone.0048358.

Mulcahy DG, Noonan BP, Moss T, Townsend TM, Reeder TW, Sites JW, Wiens JJ. Comparing the performance of dating methods using phylogenomic data in squamate reptiles. Molecular Phylogenetics & Evolution. 2012 Oct;65:974-994.

Werneck FP, Gamble T, Colli GR, Rodrigues MT, Sites JW. Deep diversification and long-term persistence in the South American ‘dry diagonal’: integrating continent-wide phylogeography and distribution modeling of geckos. Evolution. 2012 Oct;66:3014-3034.

Werneck FP, Nogueira C, Colli GR, Sites JW, Costa GC. Climatic stability in the Brazilian Cerrado: implications for biogeographical connections of the South American savannas, species richness, and conservation in a biodiversity hotspot. Journal of Biogeography. 2012 Oct;39:1695-1706.

Avile-Pires TC, Mulcahy DG, Werneck FP, Sites JW. Phylogeography of Kentropyx calcarata and Gonatodes humeralis (Reptilia: Squamata) in eastern Amazonia, Brazil. Herpetologica. 2012 Sep;68:272-287.

Wiens JJ, Hutter CR, Mulcahy DG, Noonan BP, Townsend TM, Sites JW, Reeder TW. Resolving the phylogeny of lizards and snakes with extensive sampling of genes and species. Biology Letters. 2012 Sep:doi:10.1098/rsbl.2012.0703.

Camargo A, Morando M, Avila LJ, Sites JW. Species delimitation with ABC and other coalescent-based methods in lizards of the Liolaemus darwinii complex (Squamata: Liolaemidae). Evolution. 2012 Sep;66:2834-2849.

Avila LJ, Perez CH, Medina CD, Sites JW, Morando M. A new species lizard of the Liolaemus elongatus clade (Squamata: Iguania: Liolaemini) from Curi Leuvu River Valley, northern Patagonia, Neuquén, Argentina. Zootaxa. 2012 Jun;3325:37-52.

Camargo A, Avila LJ, Morando M, Sites JW. Accuracy and precision in species tree estimation: an empirical evaluation of performance in lizards of the Liolaemus darwinii group (Squamata: Liolaemidae) under varying sub-sampling designs. Systematic Biology. 2012 Mar;61:272-288.

Sites JW. Wilmer Webster Tanner: 17 December 1909 – 28 October 2011. Herpetological Review. 2012 Mar;43:12-15.

Fontanella FM, Feltrin N, Avila LJ, Sites JW, Morando M. Early stages of divergence: phylogeography, climate modeling, and niche differentiation in the South American lizard Liolaemus petrophilus (Squamata: Liolaemidae). Ecology and Evolution. 2012 Mar;2:792-808.

Hannah N, Brown D, Avila LJ, Sites JW, Morando M, Fontanella FM. Characterization of 10 polymorphic microsatellite loci in the South American lizard Liolaemus fitsingerii with cross-amplifiction in L. chehuachekenk. Conservation Genetics Resources. 2012 Feb;4:105-107.

Mendelson JR, Mulcahy DG, Williams TS, Sites JW. A phylogeny and evolutionary natural history of Mesoamerican toads (Anura: Bufonidae: Incilius). Zootaxa. 2011 Dec;3138:1-34.

Sites JW, Avila LJ. A new species of the Liolaemus elongatus group (Squamata: Iguania: Liolaemini) from Cordillera del Veinto, northwestern Patagonia, Neuquen, Argentina. Zootaxa. 2010 Nov;2667:28-42.

Wiens JJ, Kuczynski CA, Townsend T, Reeder TW, Mulcahy D, Sites JW. Combining phylogenomics and fossils in higher-level squamate phylogeny: molecular data change the placement of fossil taxa. Systematic Biology. 2010 Oct;59(6):674-688. <website>

Camargo A, Sinervo BR, Sites JW. Lizards as model organisms for linking phylogeographic and speciation studies. Molecular Ecology. 2010 Aug;19(16):3250-3270. <website>

Sinclair EA, Pramuk J, Bezy RL, Crandall KA, Sites JW. DNA evidence for non-hybrid origins of parthenogensis in natural populations of vertebrates. Evolution. 2010 May;64(5):1346-1357.

Sinervo B, Méndez-de-la-Cruz F, Miles DB, Heulin B, Bastiaans E, Villagrán-Santa Cruz M, Lara-Resendiz R, Martínez-Méndez N, Calderón-Espinosa ML, Meza-Lázaro RN, et al. Erosion of lizard diversity by climate change and altered thermal niches. Science. 2010 May;328(5980):894-899. <website>

Avila LJ, Perez C, Morando M, Sites JW. A new species of Liolaemus (Reptilia: Squamata) from southwestern Rio Negro province, northern Patagonia, Argentina. Zootaxa. 2010 Apr;2434:47-59. <website>

Benavides E, Baum R, Snell HM, Snell HL, Sites JW. Island biogeography of Galapagos lave lizards (Tropiduridae: Microlophus): species diversity and colonization of the archipelago. Evolution. 2009 Jul;63:1606-1626.

Crother BI, Boundy J, Campbell JA, de Quieroz K, Frost D, Green DM, Highton R, Iverson JB, McDiarmid RW, Meylan PA, et al. Scientific and standard English names of amphibians and reptiles of North America north of Mexico: update. Herpetological Review. 2003 Oct;34::196-203.

Sites JW, Marshall JC. Species delimitation: a Renaissance issue in systematic biology. Trends in Ecology and Evolution. 2003;18::462-470.

Presentations

Camargo A, Morando M, Sites JW, Avila LJ. Evolución morphológico en el grupo Liolaemus darwinii (Squamata: Liolaemidae) sugiere diversificación adaptative asociada a nichos climáticos. II Congreso Uruguayo de Zoología. Montevideo, UR. 2012 Dec.

Camargo A, Werneck F, Morando M, Sites JW, Avila LJ. Expansión de rango, dinámica demográphica y paleo-distribución de Liolaemus darwinii (Squamata: Liolaemidae) durante el Cuaternario del Argentina Central. II Congreso Uruguayo de Zoología. Montevideo, UR. 2012 Dec.

Camargo A, Morando M, Sites JW, Avila LJ. Evolución morphológico en el grupo Liolaemus darwinii (Squamata: Liolaemidae) sugiere diversificación adaptative asociada a nichos climáticos. XIII Congreso Argentina Herpetologia. Mar de La Plata, ARG. 2012 Nov.

Camargo A, Werneck F, Morando M, Sites JW, Avila LJ. Expansión de rango, dinámica demográphica y paleo-distribución de Liolaemus darwinii (Squamata: Liolaemidae) durante el Cuaternario del Argentina Central. XIII Congreso Argentina Herpetologia. Mar de La Plata, ARG. 2012 Nov.

Victoriano P, Vera-Escalona I, Coronado T, Muñoz-Mendoza C, Sites JW. Tendencias geográficas de valores de conservación intraespecíficos en lagartijas del centro-sur de Chile. PIRE-Patagonia meeting: “Conservación en Patagonia: Integrando Ciencia y Gestión”. Bariloche, Argentina. 2012 Sep.

Olave M, Avila L, Sites JW, Morando M. A well-supported species tree of the Eulaemus clade (Squamata: Liolaemus): evidence of an ancient radiation. VII World Congress of Herpetology. Vancouver, BC. 2012 Aug.

Breitman MF, Avila LJ, Parra M, Sites JW, Morando M. How lizards survived blizzards: phylogeography of the Liolaemus lineomaculatus group (Liolaemini) reveals multiple breaks and refugia in southern Patagonia, and their concordance with other co-distributed taxa. VII World Congress of Herpetology. Vancouver, BC. 2012 Aug.

Morando M, Avila LJ, Perez CH, Hawkins M, Sites JW. How many Phymaturus (Squamata, Liolaemini) are there? A molecular assessment of flat lizards on rock outcrops. VII World Congress of Herpetology. Vancouver, B.C., Canada. 2012 Aug.

Werneck FP, Gamble T, Colli G, Rodrigues MT, Sites JW. Lizard diversification in the South American dry diagonal. VII World Congress of Herpetology. Vancouver, BC. 2012 Aug.

Morando M, Avila LJ, Dawson A, Perez C, Sites JW. Molecular phylogeny of Homonota (Phyllodactylidae): yet another cryptically diverse gecko clade. VII World Congress of Herpetology. Vancouver, BC. 2012 Aug.

Morando M, Avila LJ, Baker E, Sites JW. Molecular phylogeny of the endemic South American lizard clade Leiosaurae. VII World Congress of Herpetology. Vancouver, B.C., Canada. 2012 Aug.

Camargo A, Morando M, Avila LJ, Sites JW. Species delimitation with ABC and other coalescent-based methods. VII World Congress of Herpetology. Vancouver, BC. 2012 Aug.

Olave M, Avila LJ, Sites JW, Morando M. Statistical tests for an ancient radiation of the most diverse reptile genus in South America. 1st Joint Congress of Evolutionary Biology. Ottawa, Ontario, Canada. 2012 Jul.

Sites JW. Species Delimitation: A Decade After the Renaissance. invited research seminar. Brasilia, BRAZIL. 2012 Jun.

Sites JW. Species Delimitation: the Other Goal of Systematic Biology.". invited research seminar. Brasilia, BRAZIL. 2012 Jun.

Brown D, Sites JW, Fontanella FM. A multi-locus phylogenetic analysis of the North American xenodontine snakes. National Conference for Undergraduate Research. Ogden, Ut. 2012 Mar.

Baker E, Morando M, Avila LJ, Sites JW, Fontanella FM. Phylogenetic analysis of the Squamate clade Leiosauridae. National Conference for Undergraduate Research. Ogden, Ut. 2012 Mar.

Victoriano P, Gouin N, D'Elia G, Coronado T, Munoz C, Vera I, Sites JW. Comparing phylogeographies of lizards from the Chilean Biodiversity Hotspot: associations with glaciations and landscape structure. Chilean Evolution Society - annual meeting. Concepcion. 2011 Oct.

Munoz C, Gouin N, D'Elia G, Sites JW, Victoriano P. Phylogeographic comparisons of two Liolaemus from south-central Chile suggest discordant micro-evolutionary processes. Chilean Evolution Society - annual meeting. Concepcion. 2011 Oct.

Sites JW. Phylogenetic Insights on Evolutionary Novelties in Lizards and Snakes. EcoLunch. BYU-Provo. 2011 Sep.

Perez CH, Avila LJ, Olave M, Perez DR, Sites JW, Morando M. A new species of Liolaemus (Iguania: Liolaemini) from a volcanic mountain of northern Neuquen Province, Argentina. XII Congreso Argentino de Herpetología. Bariloche, Argentina. 2011 Sep.

Camargo A, Avila LJ, Morando M, Sites JW. Accuracy and precision of species trees: the effects of locus, individual, and base-pair sampling on inference of species trees in the Liolaemus darwinii group (Squamata, Liolaemidae). XII Congreso Argentino de Herpetología. Bariloche, Argentina. 2011 Sep.

Breitman MF, Parra M, Avila LJ, Sites JW, Morando M. Descrubriendo patrones genéticos y morfológicos de la Patagonia Austral: Lagartijas del grupo Lineomaculatus (Squamata, Iguania, Liolaemini, Liolaemus). XII Congreso Argentino de Herpetología. Bariloche, Argentina. 2011 Sep.

Ferraras I, Olave M, Avila LJ, Sites JW, Morando M. Estudio de la variación fenotípica en los complejos de lagartijas boulengeri y rothi (Squamata: Liolaemus). XII Congreso Argentino de Herpetología. Bariloche, Argentina. 2011 Sep.

Olave M, Avila LJ, Sites JW, Morando M. Filogenia multilocus del clado Eulaemus (Squamata: Liolaemus): Evidencias de division de linaje incomplete, hibridizacion y nuevas species candidatas. XII Congreso Argentino de Herpetología. Bariloche, Argentina. 2011 Sep.

Medina CD, Avila LJ, Perez CH, Perez DR, Sites JW, Morando M. New species of lizards of the Liolaemus elongatus species group (Iguania: Liolaemini) from northwestern Patagonia, Neuquen Province, Argentina. XII Congreso Argentino de Herpetología. Bariloche, Argentina. 2011 Sep.

Camargo A, Morando M, Avila LJ, Sites JW. Species delimitation with ABC and other coalescent-based methods of lizards in the Liolaemus darwinii complex (Squamata, Liolaemidae). XII Congreso Argentino de Herpetología. Bariloche, Argentina. 2011 Sep.

Breitman MF, Perez CH, Parra M, Sites JW. Tres nuevas species de la sección Liolaemus lineomaculatus (Squamata: Sauria: Liolaemidae) de Patagonia. XII Congreso Argentino de Herpetología. Bariloche, Argentina. 2011 Sep.

Aguilar C, Sites JW. A provisional assessment of species limits between two Andean populations of Liolaemus from the montanus group in southern Peru (Squamata; Liolaemidae). IX Latin American Congress of Herpetology. Curitiba, Brazil. 2011 Jul.

Avila-Pires LJ, Medina C, Perez C, Perez D, Sites JW, Morando M. New species of lizards previously confused with Liolaemus elongatus Koslowsky 1896 (Reptilia: Squamata: Iguania: Liolaemini) from Neuquen Province, Argentina. IX Latin American Congress of Herpetology X. Curitiba, Brazil. 2011 Jul.

Sites JW. Rock-Paper-Scissors, Viviparity, and Speciation: Links Between Climate Warming and Lizard Extinctions. Invited Seminar Speaker. Department of Biology, Weber State University. 2011 Apr.

Hannah N, Morando M, Avila LJ, Sites JW, Fontanella F. A microsatellite library of the South American genus Liolaemus: provisional tests of their utility for evolutionary studies. College of Life Sciences: Undergraduate Research Symposium. BYU-Provo. 2011 Mar.

Brown D, Sites JW, Fontanella F. Multi-locus phylogenetic analysis of the North American xenodontine snakes. College of Life Sciences: Undergraduate Research Symposium. BYU-Provo. 2011 Mar.

Baker E, Morando M, Avila LJ, Sites JW, Fontanella F. New insights into phylogenetic relationships and species diversity of the South American lizard species complex Liolaemus fitzingerii, based on combined analysis of mitochondrial and nuclear data. College of Life Sciences: Undergraduate Research Symposium. BYU-Provo. 2011 Mar.

Sites JW. Rock-Paper-Scissors, Viviparity, and Speciation: Links Between Climate Warming and Lizard Extinctions. Cross-program Seminar Speaker. Center for the Environment/Discovery Center, Purdue University. 2011 Mar.

Sites JW. Rock-Paper-Scissors, Viviparity, and Speciation: Links Between Climate Warming and Lizard Extinctions. EcoLunch. BYU-Provo. 2011 Mar.

Technical Report

Minoli I, Morando M, Breitman F, Medina C, Sites JW, Avila LJ. "Lagartijas de la Provincia de Chubut (Argentina): Distribución Geográfica, Diversidad Genética y Comentarios Acerca de su Estado de Conservación. Informe Técnico .". 2012 Oct.