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Ryan Stewart

Plant & Wildlife Sciences

2124 LSB
Provo, UT 84602


  • Ph.D., Horticulture, Iowa State University, 2005
  • M.S., Plant Science, Utah State University, 2001
  • B.S., Agricultural Systems Technology, Utah State University, 1999



  • Professor, Brigham Young University, 2018-Present
  • Associate Professor, Brigham Young University, 2011-2018
  • Associate Professor, University of Illinois, 2011-2011
  • Assistant Professor, University of Illinois, 2005-2011

Honors and Awards

  • Hokkaido University : Visiting Associate Professor Fellowship
  • University of Illinois : Teaching Scholar
  • University of Illinois : Global Academy Fellow
  • University of Illinois : Global Academy Scholar

Courses Taught

Winter 2019

  • PWS 301: Plant Growth & Reproduction Section 001, 002
  • PWS 799R: Doctoral Dissertation Section 004

Fall 2018

  • PWS 191: Intro to Landscape Management Section 001
  • PWS 210: Herbaceous Plants Section 001

Spring 2018

  • PWS 799R: Doctoral Dissertation Section 007

Winter 2018

  • PWS 100: Plants in the Environment Section 001
  • PWS 301: Plant Growth & Reproduction Section 001, 002


Journal Articles

Dong H, Green SV, Nishiwaki A, Yamada T, Stewart JR, Deuter M, Sacks EJ. 2018. Winter hardiness of Miscanthus (I): Overwintering ability and yield of new Miscanthus ×giganteus genotypes in Illinois and Arkansas. Global Change Biology Bioenergy.

Brumfield RG, Kenny LB, DeVincentis AJ, Koeser AK, Verlinden S, Both AJ, Bi G, Lovell ST, Stewart JR. 2018. Analysis of economic and social costs of growing Petunia xhybrida in a greenhouse production system using alternative containers. HortScience. 53:1179-1185.

Toma Y, Yamada T, Fernández FG, Nishiwaki A, Hatano R, Stewart JR. 2016. Evaluation of greenhouse gas emissions in a Miscanthus sinensis dominated semi-natural grassland in Kumamoto, Japan. Soil Science and Plant Nutrition. 62:80-89.

Bergsten SJ, Koeser AK, Stewart JR. 2016. Evaluation of the impacts of salinity on biomass and nutrient levels of Agave species with agricultural potential in semiarid regions. HortScience. 51:1-6.

Stewart JR. 2015. Agave as a model CAM crop system for a warming and drying world. Frontiers in Plant Science.

Yang X, Cushman JC, Borland AM, Edwards EJ, Wullschleger SD, Tuskan GA, Owen NA, Griffiths H, Smith JA, De Paoli HC, et al. 2015. A roadmap for research on crassulacean acid metabolism (CAM) to enhance sustainable food and bioenergy production in a hotter, drier world. New Phytologist. 207:491-504.

Sun Y, Niu G, Koeser AK, Bi G, Anderson V, Jacobsen K, Conneway R, Verlinden S, Stewart JR, Lovell ST. 2015. Impact of biocontainers on plant performance and container decomposition in the landscape. HortTechnology. 25:35-41.

Evans MR, Koeser AK, Bi G, Nambuthiri S, Geneve R, Lovell ST, Stewart JR. 2015. Impact of biocontainers with and without shuttle trays on water use in the production of a containerized ornamental greenhouse crop. HortTechnology. 25:35-41.

Conneway R, Verlinden S, Koeser AK, Evans M, Schnelle R, Anderson V, Stewart JR. 2015. Use of alternative containers for long- and short-term greenhouse crop production. HortScience. 25:26-34.

Brumfield RG, DeVincentis AJ, Wang X, Fernandez RT, Nambuthiri S, Geneve RL, Koeser AK, Bi G, Li T, Sun Y, et al. 2015. Economics of utilizing alternative containers in ornamental crop production systems. HortTechnology. 25:17-25.

Clark LV, Stewart JR, Nishiwaki A, Toma Y, Kjeldsen JB, Jorgensen U, Zhao H, Peng J, Yoo JH, Heo K, et al. 2015. Genetic structure of Miscanthus sinensis and M. sacchariflorus in Japan indicates a gradient of bidirectional but asymmetric introgression. Journal of Experimental Botany. 66:4213-4225.

Nadir M, Tanaka A, Kuwabara S, Matuura H, Yamada T, Stewart JR, Nishiwaki A. 2014. Comparison of relative DNA content estimated using DAPI and PI-FCM in Miscanthus sinensis, Miscanthus sacchariflorus, and their hybrids. Journal of Warm Regional Society of Animal Science (日本暖地畜産学会報). 57:53-57.

Dwiyanti MS, Stewart JR, Yamada T. 2014. Forages for feedstocks of biorefineries in temperate environments: review of lignin research in bioenergy crops and some insight into Miscanthus studies. Crop and Pasture Science. 65:1199-1206.

Dwiyanti MS, Stewart JR, Nishiwaki A, Yamada T. 2014. Natural variation in Miscanthus sinensis seed germination under low temperatures. Grassland Science. 60:194-198.

Byers C, Maughan PJ, Clouse J, Stewart JR. 2014. Microsatellite primers in Agave utahensis (Agavaceae), a keystone species in the Mojave Desert and Colorado Plateau. Applications in Plant Sciences.

Głowacka K, Clark LV, Adhikari S, Peng J, Stewart JR, Nishiwaki A, Yamada T, Jorgensen U, Hodkinson TR, Gifford J, et al. 2014. Genetic variation in Miscanthus ×giganteus and the importance of estimating genetic distance thresholds for differentiating clones. Global Change Biology Bioenergy. 7:386-404.

Bergsten SJ, Stewart JR. 2014. Measurement of the influence of low water availability on the productivity of Agave weberi, cultivated under controlled irrigation. Canadian Journal of Plant Science. 94:439-444.

Koeser AK, Lovell ST, Petri AC, Brumfield RG, Stewart JR. 2014. Biocontainer use in Petunia xhybrida greenhouse production – a cradle-to-gate carbon footprint assessment of secondary impacts. HortScience.

Koeser AK, Lovell ST, Evans MR, Stewart JR. 2013. Biocontainer water use in short-term greenhouse crop production. HortScience. 23:215-219.

Dwiyanti M, Rudolph A, Swaminathan K, Nishiwaki A, Shimono Y, Kuwabara S, Matuura H, Nadir M, Moose S, Stewart JR, et al. 2013. Genetic analysis of putative triploid Miscanthus hybrids and tetraploid M. sacchariflorus collected from sympatric populations of Kushima, Japan. BioEnergy Research. 6:486-493.

Howlett DS, Toma Y, Wang H, Sugiyama S, Yamada T, Nishiwaki A, Fernandez F, Stewart JR. 2013. Soil carbon source and accumulation over 12,000 years in a semi-natural Miscanthus sinensis grassland in southern Japan. Catena. 104:127-135.

Toma Y, Armstrong K, Stewart JR, Yamada T, Nishiwaki A, Fernandez F. 2012. Carbon sequestration in soil in a semi-natural Miscanthus sinensis grassland and Cryptomeria japonica forest plantation in Aso, Kumamoto, Japan. Global Change Biology Bioenergy. 4:566-575.

Matlaga DP, Quinn LD, Davis AS, Stewart JR. 2012. Light response of native and introduced Miscanthus sinensis seedlings. Invasive Plant Science and Management. 5(3):363-374.

Quinn L, Culley T, Stewart JR. 2012. Genetic comparison of introduced and native populations of Miscanthus sinensis (Poaceae), a potential bioenergy crop. Grassland Science. 58(2):101-111.

Quinn L, Stewart JR, Yamada T, Toma Y, Saito M, Shimoda K, Fernandez F. 2012. Environmental tolerances of Miscanthus sinensis in invasive and native populations. BioEnergy Research. 5:139-148.

Toma Y, Sato S, Izumi M, Fernandez FG, Stewart JR, Hatano R, Nishiwaki A, Yamada T. 2012. Response of fertilization to aboveground biomass production of Miscanthus sinenisis of poor fertility soil – Case study in Tomakomai, Hokkaido. Research Bulletin of the Hokkaido National Agricultural Experiment Station. 79:162-169.

Fedewa CA, Stewart JR. 2011. Field establishment of little bluestem and prairie dropseed under managed conditions. Native Plants Journal. 12:111-117.

Bayer AL, Stewart JR. 2011. Prospects for conservation of an endemic woody species native to Florida, Chionanthus pygmaeus (pygmy fringetree), through seed and vegetative propagation. Native Plants Journal. 12:62-69.

Toma Y, Fernández FG, Sato S, Izumi M, Hatano R, Yamada T, Nishiwaki A, Bollero G, Stewart JR. 2011. Carbon budget and methane and nitrous oxide emissions over the growing season in a Miscanthus sinensis grassland in Tomakomai, Hokkaido, Japan. Global Change Biology Bioenergy. 3:116-134.

Nishiwaki A, Mizuguti A, Kuwabara S, Toma Y, Ishigaki G, Miyashita T, Yamada T, Matuura H, Yamaguchi S, Rayburn AL, et al. 2011. Discovery of natural Miscanthus (Poaceae) triploid plants in sympatric populations of Miscanthus sacchariflorus and Miscanthus sinensis in southern Japan. American Journal of Botany. 98:154-159.

Quinn LD, Matlaga DP, Stewart JR, Davis AS. 2011. Empirical evidence of long-distance dispersal in Miscanthus sinensis and Miscanthus x giganteus. Invasive Plant Science and Management. 4:142-150.


Stewart JR. Agave, a drought-tolerant succulent crop, enabled ancient tribes to survive and could allow modern economies to thrive in the semi-arid southwestern U.S. Hokkaido University Sustainability Week Conference – How can ancient agricultural practices in Japan and the USA inform modern agriculture sustainability?. Sapporo, Japan. 2017.

Stewart JR. Exploring the possibility of photosynthetic plasticity in Agave sensu lato. IX International Congress on Cactus Pear and Cochineal – CAM crops for a hotter and drier world. Coquimbo, Chile. 2017.

Anderson B, Evans M, Wallace L, Clouse J, Maughan PJ, Stewart JR. SNP-based Genetic Diversity and Structure of Agave utahensis. Botany 2016. Savannah, Georgia. 2016.